A
B C D
E F G H
I J K L
M N O P
Q R S T
U V W X Y
Z
Abaxial. Away from the axis; the
lower surface of the leaf; dorsal
Abcissent. Falling off or separating at a specific
separation (abcission) layer, as in most deciduous plants
Acarpous. No carpels or carpellate whorl; no pistil
Accessory Bud. Buds lateral to or above axillary buds; Accessory
Organs - The calyx and corolla
Accrescent. Growing after flowering or bud development
has occurred, as the sepals in Hypericum and bud scales in Carya
Accumbent or Pleurorhizal. Reclinate with cotyledon
edges against hypocotyl
Acerose. Needle-shaped; sharp
Achene. A one-seeded, dry, indehiscent fruit with
seed attached to fruit wall at one point only, derived from
a one-loculed superior ovary
Achenecetum. An aggregation of achenes, as in
Ranunculus
Achlamydeous. Without perianth
Acicular. Needlelike, round or grooved in cross
section
Aciculate . Finely marked as with pin pricks,
fine lines usually randomly arranged
Acrocaulous. With terminal branches
Acrocidal Capsule. One that dehisces through terminal
slits, or fissures, as in Staphylea
Acrodromous. With two or more primary or strongly
developed secondary veins diverging at or above the base of
the blade and running in convergent arches toward the apex over
some or all of the blade length, the arches not basally curved
Acropetal. Developing upward, toward apex
Acroramous. Leaves terminal, near apex of branch
Acroscopic. Facing apically
Actinodromous. With three or more primary veins
diverging radially from a single point at or above the base
of the blade and running toward the margin, reaching it or not
Actinomorphic or Radial. With floral parts radiate
from center like spokes on wheel
Actinostele. A protostele having a xylem core
in the form of radiating ribs, as viewed in transverse section
Aculeate. Prickly
Acuminate (Base narrowly cuneate). Margins straight
to convex forming a terminal angle of less than 45
Acute (Base cuneate). Margins straight to convex
forming a terminal angle 45-90
Adaxial. Next to the axis; facing the stem; ventral
Adherent. With unlike parts of organs joined, but only
superficially and without actual histological continuity
Adnate. With unlike parts or organs integrally
fused to one another with histological continuity
Adventitious. Arising from organ other than root;
usually lateral
Aerating
or Knee. Vertical or horizontal aboveground
roots
Aerial
or Epigeous. Above the ground or water, in the
air; aerial stem - an erect stem arising from a horizontal rhizome
Aerocaulous. With aerial stems
Aerophyllous. With aerial leaves
Aestival. Appearing in summer
Agamandrous. Inflorescence with neuter flowers
inside or above and staminate outside or below (agamandrocephalous)
Agamogynous. Inflorescence with neuter flowers
inside or above and pistillate outside or below (agamogynecephalous)
Agamohermaphroditic. Inflorescence with neuter
flowers inside or above and hermaphroditic outside or below
(agamohermaphrodicephalous)
Agamous
or Neuter. Without sex; sexual organs abortive
Agglomerate, Conglomerate, Crowded, or Aggregate.
Dense structures with varied angles of divergence
Aianthous or Semperflorous. With flowers appearing
throughout the year
Alate. Winged
Allagostemonous. Having stamens attached to petal
and torus alternately
Allautogamy. Cross- and self-fertilization in
same plant, as in Viola
Allogamy
or Xenogamy. Cross-fertilization in plants
Allopatric. Isolated, separated
Alternate. One leaf or other structure per node
Alveolate. Honey-combed
Ament
or Catkin. A unisexual spike or elongate axis
with simple dichasia that falls as a unit after flowering or
fruiting
Amorphic
or Paleomorphic. Flowers without symmetry; usually
with an indefinite number of stamens and carpels, and usually
subtended by bracts or discolored upper leaves; e.g., Salix
discolor, Echinops ritro (mostly fossil forms)
Amphicarpous. With fruits in two environments;
e.g., aerial and subterranean
Amphiflorous. Flowers above and below ground,
as in Amphicarpum
Amphisarca. A berry-like succulent fruit with
a crustaceous or woody rind, as in Lagenaria
Amphitropous. With body bent or curved on both
sides so that the micropyle is near the medially attached funiculus
Amplexicaul. Completely clasping the stem
Ampliate. Enlarged; dilated
Anadromous. Having the first lobe or segment of
a pinna arising basiscopically in compound leaves
Anatropous. With body completely inverted so that
funiculus is attached basally near adjoining micropyle area
Ancipital. Two-edged
Andragamous. Inflorescence with staminate flowers
inside or above and neuter flowers outside or below (andragamocephalous)
Androdioecious. Some plants with staminate flowers
and some with perfect flowers
Androecium. One or more whorls or groups of stamens;
all stamens in flower
Androgynecandrous. Inflorescence with staminate
flowers above and below pistillate, as in the spikes of some
species of Carex
Androgynophore. The stipe or column on which stamens
and carpels are borne
Androgynous. Inflorescence with staminate flowers
inside or above and pistillate outside or below (androgynecephalous)
Androhermaphroditic. Inflorescence with staminate
flowers inside or above and hermaphroditic outside or below
(androhermaphrodicephalous)
Andromonoecious. Plant with staminate and perfect
flowers
Anemophily. Pollinated by wind
Angustate. Narrow
Anisocarpous. With unequal carpels
Anisocotylous. With unequal cotyledons
Anisolateral. With unequal sides
Anisopetalous. With unequal petals
Anisophyllous. With unequal leaves
Anisostylous. With unequal styles
Annotinal or Yearly. Appearing yearly
Annual. Living one year or less. Winter Annual
- Living less than one year but through the winter; germination
usually in late fall, and usually flowering and fruiting in
early spring
Annular. Ring-like
Annulus. Thick-walled ring of cells on the sporangium
Anomalicidal or Rupturing Capsule. One that dehisces
irregularly, as in Ammannia
Anterior. Anterior Lobes - The lobes away from
axis, toward the subtending bract, abaxial lobes; Anterior Ridges,
Lines, Grooves - The lines, grooves, ridges in or on the dorsal
side, abaxial, within the perianth
Anther. Pollen-bearing portion of stamen
Antheridium. The male sex organ producing the
sperm
Anthesis. Time of flowering; opening of flower
with parts available for pollination
Anthocarpous. Having a body of combined floral
and fruit parts, as in multiple fruits
Anthotaxis. Arrangement of sporophylls, primarily
reproductive in function
Anthropophily. Pollinated by man
Antipetalous. Opposite the petals
Antisepalous. Opposite the sepals
Antitropous. With radicle pointing away from hilum
Antrorse. Bent or directed upward
Apetalous. No petals or corolla
Aphyllopodic. Without blade-bearing leaves at
base of plant
Aphyllous. Without leaves, no whorls of leaves
Apical
or Terminal. At the top, tip, or end of a structure
Apiculate. More than 3:1 l/w, usually slightly
curled and flexuous
Apocarpous. With carpels separate
Apogamy. Producing sporophytes from a gametophyte
without fertilization
Apopetalous or Choripetalous. With separate petals
Apophysis. Exposed outer surface of either an
ovuliferous scale or megasporophyll as seen when the cone is
closed
Aposepalous or Chorisepalous. With separate sepals
Apospory. Producing gametophytes directly from
a sporophyte without producing spores
Apostemonous. With separate stamens
Appendicular. Typical stamen with a variously-shaped
or modified, protruding connective, as in Viola
Applanate or Plane. Flat, without vertical curves
or bends
Appressed or Adpressed. Pressed closely to axis
upward with angle of divergence 15° or less
Arachnoid. Cobwebby
Arborescent. Tree-like in appearance and size
Archegonium. The female sex organ producing the
egg
Arcuate. Curved like a crescent, can be downward
or upward
Areolate. Divided into many angular or squarish
spaces
Areoles. The spaces formed by a vein network
Arhizous. Without roots, no whorls of roots
Aril. Outgrowth of funiculus, raphe, or integuments;
or fleshy integuments or seed coat, a sarcotesta at base of
the fleshy seed; e.g., Cepalotaxus
Arillate. General term for an outgrowth from the
funiculus, seed coat or chalaza; or a fleshy seed coat
Aristate. More than 3:1 l/w, usually prolonged,
straight and stiff
Articulate. Generally meaning having a joint as
in leaves, leaflets or stems, as in heterophyllous species of
Selaginella; or having a swollen area, often discolored, at
the point of branching of the stem
Ascending. Directed upward with an angle of divergence
of 16-45°
Asepalous. No sepals or calyx
Asperous. Having a rough surface
Assurgent. Directed upward or forward
Astemonous or Anandrous. No stamens or androecium
Astylocarpellous. Without a style and a stipe
Astylocarpepodic. Without a style, with a stipe
Astylous. Style absent
Asymmetric. Without regularity in any dimension
Attenuate. Elongate, tapering, usually applied
to base
Auriculate. Lobe rounded; sinus depth variable;
outer margin concave, inner convex or straight
Auriculiform. Usually obovate with two small rounded,
basal lobes
Autogamy. Self-fertilization in a single flower
Autumnal. Appearing in autumn
Awl-shaped Leaf. Subulate; narrow, flat, stiff,
sharp-pointed, usually less than 1/2 in. long; e.g., Juniperus
Axial. With branches arising from buds in leaf
axil
Axile. With the placentae along the central axis
in a compound ovary with septa
Axillary
or Lateral. In axils of leaves or leaf scars;
axillary leaves - leaves borne in the axils of branches, as
in heterosporous species of Selaginella Baccacetum or Etaerio.
An aggregation of berries, as in Actaea
Baccate. Juicy and very succulent
Balausta. Many-seeded, many- loculed indehiscent
fruit with a tough, leathery pericarp, as in Punica
Banded. Transverse stripes of one color crossing
another
Barbed. With short, rigid reflexed bristles or
processes
Barbellate. Minutely barbed
Bark. Tissues of plant outside wood or xylem
Basal
or Radical. At the bottom or base of a structure
Base. bottom or lower portion
Basicaulous. With basal branches
Basicaulous. Near base of stem
Basicidal Capsule. One that dehisces through basal
slits or fissures, as in some species of Aristolochia
Basifixed. Anther attached at its base to apex
of filament
Basipetal. Developing downward, toward base
Basipetiolar. At the base of the petiole
Basiramous. Leaves on lower part of branch
Basiscopic. Facing basally
Bast
bundles (peripheral strands). bundles of thick-walled
cells parallel to the midrib, as in Isoetes
beard. A tuft, line or zone of trichomes
Bearded
or Barbate. With long trichomes usually in a
tuft, line or zone
Bent. Foliate embryo with expanded and usually
thick cotyledons in an axile position bent upon the hypocotyl
in a jacknife position
Berry. Fleshy fruit, with succulent pericarp,
as in Vitis
Bibacca. A fused double berry, as in Lonicera
Bicarpellate. Two-carpelled
Bicrenate or Doubly-crenate. With smaller rounded
teeth on larger rounded teeth
Bidentate. Two-toothed
Biduous. Lasting two days
Biennial. Living two years, usually flowering
second year
Biferous. Appearing twice yearly
Bifid. Cut or divided into two lobes or parts
Biflorous. Flowering in autumn as well as in spring
Bifoliolate, Geminate, or Jugate. With two leaflets
from a common point
Bifurcate. Divided into two forks or branches
Bigeminate, Bijugate. With two orders of leaflets,
each bifoliolate; doubly paired
Bilabiate. Two-lipped, with two unequal divisions
Bilocular. Two-locular
Bimestrial. Periodicity: occurring every two months;
duration: lasting two months
Binate. Twinned
Bipalmate. Twice palmate
Bipinnate. Twice pinnate
Biseriate. Two-rowed; in two series
Biserrate or Doubly-serrate. With sharply cut
teeth on the margins of larger sharply cut teeth
Bisexual. Both sexes in same flower (monoclinous,
perfect)
Biternate. With two orders of leaflets, each ternately
compound
Blade. The expanded portion of a leaf
Blastocarpous. Germination of seeds while within
the pericarp, as in Rhizophora
Blotched. The color disposed in broad, irregular
blotches
Bordered. One color is surrounded by an edging
of another
Botuliform. Sausage-shaped
Bract. Modified, usually reduced, leaf in the
inflorescence
Bracteal
and Laminar. May be localized or found over
entire structure
Bracteole or Bractlet. A secondary or smaller
bract
Bristle. A stiff, strong trichome, as in the perianth
of some members of the Cyperaceae
Bristly. Beset with bristles
Brochidodromous. With a single primary vein, the
secondary veins not terminating at the margin but joined together
in a series of prominent upward arches or marginal loops on
each side of the primary vein
Bud. Immature vegetative or floral shoot or both,
often covered by scales. Bud Primordium - Meristematic tissue
that gives rise to a lateral bud
Bulb. A short, erect, underground stem surrounded
by fleshy leaves
Bulbel. A small bulb produced from the base of
a larger bulb
Bulbil. A small bulb or bulb-like body produced
on above ground parts
Bulblet. A small bulb, irrespective of origin;
a small, bud-like vegetative propagule produced on the leaves
of some ferns
Bullate. Puckered or blistered
Bur
(Involucre). Cypsela enclosed in dry involucre,
as in Xanthium
Buttress. Roots with board-like or plank-like
growth on upper side, presumably a supporting structure
Caducous. Dropping off very early,
usually applied to floral parts
Calcarate. Spurred
Calceolate. Slipper-shaped, as in the corolla
of Cypripedium
Callosity. A thickened, raised area, which is
usually hard; a callus
Calybium. A hard one-loculed dry fruit derived
from an inferior ovary, as in Quercus
Calyx. The lowermost whorl of modified leaves,
sepals
Campanulate. Bell-shaped; with flaring tube about
as broad as long and a flaring limb
Campylodromous. With several primary veins or
their branches diverging at or close to a single point and running
in strongly developed, basally recurved arches which converge
toward the apex, reaching it or not
Campylotropous. With body bent or curved on one
side so that micropyle is near medially attached funiculus
Canaliculate. Longitudinally grooved, usually
in relation to petioles or midribs
Cancellate or Clathrate. Latticed
Canescent or Incanous. Covered with dense, fine
grayish-white trichomes
Cantharophily. Pollinated by beetles
Capillate. Hair-shapedCapitate. Head-like
Capitulum or Head. A determinate or indeterminate
crowded group of sessile or subsessile flowers on a compound
receptacle or torus
Carina. Keel. Carinal Canal - A canal beneath
a stem ridge associated with a vascular bundle
Carinate. Keeled
Carnose
or Sarcous. Fleshy
Carpel. The female sporophyll within flower; floral
organ that bears ovules in angiosperms; unit of compound pistil
Carpophore. Floral axis extension between adjacent
carpels, as in the Apiaceae
Carpopodium. Short, thick, pistillate stalk
Carpotaxis. Arrangement of fruits, reproductive
in function
Cartilaginous. Hard and tough but flexible
Carunculate. With an excrescent outgrowth from
integuments near the hilum, as in Euphorbia; fibrous with stringy
or cord-like seed coat, as mace in Myristica
Caryopsis or Grain. A one-seeded dry, indehiscent
fruit with the seed coat adnate to the fruit wall, derived from
a one-loculed superior ovary
Catadromous. Having the first lobe or segment
of a pinna arising acroscopically in compound leaves
Cataphyll. Rudimentary scale leaf produced by
seedling, usually in cryptocotylar species
Caudate
(Base attenuate). Acuminate with concave margins
Caudex. A short, thick, vertical or branched perennial
stem usually subterranean, or at ground level
Cauline. May be all over (general) or along the
ribs (costal), or in the grooves (canaliculate)
Caulocarpic. Plants having the stem living for
many years, bearing flowers and fruits
Caulous. With branches more or less evenly spaced
along trunk
Central Canal. The large centrally located
air space in the stem
Centrifugal. Developing from the inside outward,
or from top downward
Centripetal. Developing from the outside inward,
or from bottom upward
Centroramous. At the center of the branch
Ceraceous. Waxy
Cernuous. Drooping
Cespitose. Short, much-branched, plant forming
a cushion
Chaff
or Pale. Scale or bract at base of tubular flower
in composites
Chalaza. End of ovule opposite micropyle
Chalazogamy. Pollen tube entrance through chalaza
Channelled or Canaliculate. With a longitudinal
groove
Chartaceous. Papery, opaque and thin
Chasmantheric Pollination. Pollen transferred
from a normally dehisced anther by a pollinating agent with
pollen grain germination on the stigma and subsequent growth
of the pollen tube through stigma, style and the ovule into
the embryo sac
Cheiropterophily. Pollinated by bats
Chlamydeous. With perianth
Ciliate. With conspicuous marginal trichomes
Ciliolate. With tiny or small marginal trichomes
Cincinnus. A tight, modified helicoid cyme in
which pedicels are short on the developed side
Circinate. With lamina rolled from apex to base
with apex in center of coil
Circumalate. Winged circumferentially
Circumferential. At or near the circumference;
surrounding a rounded structure
Circumscissle Capsule or Pyxis. One that dehisces
circumferentially, as in Plantago
Cirrhous. More than 10:1 l/w, coiled and flexuous
Cladode
(phylloclad). A flattened main stem resembling
a leaf
Cladodromous. With a single primary vein, the
secondary veins not terminating at the margin and freely ramified
toward it
Cladoptosic. Shedding of branches, stems and leaves
simultaneously, as in Taxodium
Clambering. Sprawling across objects, without
climbing structures
Clasping. Partly surrounding the stem.
Clavate. Club-shaped
Claw. The long, narrow petiole-like base of a
sepal or petal
Cleft. Indentations or incisions cut 1/4-1/2 distance
to midrib or midvein
Cleistantheric Pollination. Pollen not transferred
from a normally dehisced anther by a pollinating agent; pollen
grain germinates within the anther with subsequent growth of
the pollen tube through the anther wall and ovary wall into
the ovule and embryo sac
Climbing. Growing upward by means of tendrils,
petioles, or adventitious roots
Clinanthium. The compound receptacle of the composite
head
Clouded. Colors are unequally blended together
Clustered, Conglomerate, Agglomerate, Crowded, Aggregate.
Parts dense, usually irregularly overlapping each other
Coalescent. With like or unlike parts or organs
incompletely separated; partially fused in a more or less irregular
fashion
Cochleate. Snail-shaped
Coenocarpium (Various Structures). Multiple fruit
derived from ovaries, floral parts, and receptacles of many
coalesced flowers, as in Ananas
Coetaneous. Flowering as the leaves expand; synantherous
Coherent. With like parts or organs joined, but
only superficially and without actual histological continuity
Coleoptile. Protective sheath around epicotyl
in grasses
Coleorhiza. Protective sheath around radicle in
grasses
Collet. External demarcation between hypocotyl
and root
Columella. The sterile central part within a mature
sporangium or capsule
Column,
Gynostemium or Gynandrium. With fused stamens
and carpels (stigma and style) as in Orchis
Columnar. Erect with a stout main stem or trunk
Comose. With a tuft of trichomes, usually apical
Complete. Leaf with blade, petiole, and stipules;
flower with four types of floral parts
Compound. Composed of two or more anatomically
or morphologically equivalent units, whether subdivided into
them or an aggregate of them
Compressed or Complanate. Flattened
Conduplicate. Longitudinally folded upward or
downward along the central axis so that ventral and/or dorsal
sides face each other
Cone
(strobilus). Aggregation of sporangia-bearing
structures at tip of the stem (either sporophylls or scales
in the Gymnosperms)
Conical. Having figure of true cone
Conjugate. Fused pairs, as the fruits of Lonicera
Connate. With like parts or organs integrally
fused to one another with histological continuity
Connective. Filament extension between thecae
Connivent. Convergent apically without fusion
Contiguous. Touching but not adnate, connate,
adherent, or coherent
Continuous. Symmetry of arrangement even, not
broken
Contorted. Twisted around a central axis; twisted
Contortuplicate. With weirdly folded corrugate
cotyledons
Contractile or Pull. Roots capable of shortening,
usually drawing the plant or plant part deeper into the soil,
usually with a wrinkled surface
Convolute. With one lamina enrolled in another
lamina
Cordate (Apex obcordate). Lobe rounded;
sinus depth 1/8-1/4 distance to midpoint of blade; margins convex
and/or straight
Cordiform. Heart-shaped
Coriaceous. Thick and leathery
Corm. The enlarged, solid, fleshy base of a stem
with scales; an upright underground storage stem
Cormel. Small corm produced at base of parent
corm
Corneous. Horny
Corniculate. Horned
Corolla. The whorl of petals located above the
sepals
Corona. A crown; any outgrowth between the stamens
and corolla which may be petaline or staminal in origin
Coronate. Tubular or flaring perianth or staminal
outgrowth; petaloid appendage
Coroniform. Crown-shaped
Corrugate. With lamina irregularly folded in all
directions, wrinkled
Corymb. A flat-topped or convex indeterminate
cluster of flowers. Compound Corymb - A branched corymb
Costa. The midvein of a minor divsion of a fern
leaf
Costate. Coarsely ribbed
Cotyledon. Embryonic leaf or leaves in seed
Cotylespermous. With food reserve in cotyledon,
derived from zygote
Cotyliform. Cup-shaped
Craspedodromous. Simple - With a single primary
vein, all of the secondary veins and their branches terminating
at the margin. Mixed - With a single primary vein, some of the
secondary veins terminating at the margin and an approximately
equal number otherwise
Crateriform. Shallow cup-shaped as the involucre
of some species of Quercus
Crenate. Shallowly ascending round-toothed, or
teeth obtuse; teeth cut less than 1/8 way to midrib or midvein
Crenulate. Diminutive of crenate, teeth cut to
1/16 distance to midrib or midvein
Crested
or Cristate. With a terminal ridge or tuft
Crispate. Curled; margins divided and twisted
in more than one plane
Crozier. The coiled developing leaf of a fern
Cruciform or Cruciate. Cross-shaped
Crustaceous. Hard, thin, and brittle
Cryptantherous. With stamens included
Cryptocotylar or Hypogeous. With the cotyledons
remaining inside the seed; seed usually remaining below ground
Cucullate. Hooded
Culm. Flowering and fruiting stems of grasses
and sedges
Cupule. Fused involucral bracts subtending flower,
as in Quercus
Curvative or arcuate. With lamina folded transversely
into an arc
Cuspidate. Acute but coriaceous and stiff
Cyathium. A pseudanthium subtended by an involucre,
frequently with petaloid glands, as in Euphorbia
Cylindric. Long-tubular
Cymbiform. Boat-shaped
Cyme
or Dichasium. A determinate, dichotomous inflorescence
with the pedicels of equal length. Scorpioid Cyme or Rhipidium
- A zigzag determinate inflorescence with branches developed
on opposite sides of the rachis alternately. Compound
Cyme
- A branched cyme
Cymule. A simple, small dichasium
Cypsela. An achene derived from a one-loculed,
inferior ovary
Deciduous. Persistent for one
growing season
Declinate. Directed or curved downward
Decompound. A general term for leaflets in two
or more orders - bi-, tri-etc - pinnately, palmately, or ternately
compound
Decumbent. Reclining or lying on the ground with
the tips ascending
Decurrent. Elongate, extending downward
Decussate. Opposite leaves at right angle to preceding
pair
Deflexed. Bent abruptly downward
Dehiscent. Opening regularly by valves, slits,
etc., as a capsule or anther
Deliquescent. Softening and wasting away
Dentate. Margins with rounded or sharp, coarse
teeth that point outwards at right angles to midrib or midvein,
cut 1/16 to 1/8 distance to midrib or midvein
Denticidal Capsule. One that dehisces apically,
leaving a ring of teeth, as in Cerastium
Denticulate. Diminutive of dentate, cut to 1/16
distance to midrib or midvein
Depauperate. Small and usually poorly developed
Depressed. Pressed closely to axis downward with
angle of divergence of 166-180°
Descending. Directed downward with an angle of
divergence of 136-165°
Determinate. Growth of plant parts, the size of
which is limited by cessation of meristematic activity during
the year
Dextrorse. Rising helically from right to left,
a characteristic of twining stems
Diadelphous. With two groups of stamens connate
by their filaments
Diandrous. With two stamens per flower
Diaphanous. Translucent
Dichasium. Cymose inflorescence in which each
axis produces a pair of lateral axes
Dichlamydeous. With perianth composed of distinct
calyx and corolla
Dichogamous. With maturation of stamens or anther
and carpels or stigma at different times
Dichotomous. With branches forking into two more
or less equal parts
Diclesium (Calyx). Achene or nut surrounded by
a persistent calyx, as in Mirabalis
Diclinous. Plant with imperfect flowers; stamens
and carpels in separate flowers either monoecious or dioecious
Dicotyledonous. With two cotyledons
Dictyostele. A dissected solenostele with each
individual bundle a meristele
Dicyclic. Two-whorled
Didymous. With stamens in two equal pairs
Didynamous. With stamens in two unequal pairs
Diffuse. Spread over a wide surface
Digestive Glands. Enzyme-secreting glands found
mostly on leaves of carnivorous and insectivorous plants
Dilated. Widened; expanded
Dimerous. Whorl with two members
Dimidiate. Divided into unequal halves
Dimorphic. Having two different sizes and/or shapes
within the same species
Dioecious. Plant with all flowers imperfect, but
staminate and pistillate on separate plants
Diplecolobal. With incumbent cotyledons folded
two or more times
Diplostemonous. With stamens in two whorls, outer
opposite the sepals, inner opposite petals
Diplotegium. A pyxis derived from an inferior
ovary
Dipterous. Two-winged
Disc. A discoid structure developed from receptacle
at base of ovary or from stamens around the ovary
Discoid. Orbicular with convex faces
Discoidal. A single large spot of color in the
center of another
Discontinuous. Basal and lateral, basal and terminal,
or lateral and terminal; not continuous
Dissected. Irregularly cut into numerous segments
Distal. Away from the point of origin or attachment
Distichous. Leaves 2-ranked, in one plane
Distinct. With like parts or organs unjoined and
separate from one another
Diurnal. Opening during the day
Divergent, Patent, or Divaricate. More or less
horizontally spreading with angle of divergence of 15° or less
up or down from the horizontal
Divided. Cut 3/4 to almost entire distance to
middle of structure
Dolabriform. Axe-shaped
Dorsal. Pertaining to the surface most distant
from the axis; back of an outer face of organ; lower side of
leaf; abaxial. Dorsal Side - Back or abaxial side, or the lower
side of a perianth part
Dorsifixed. Anther attached dorsally and medially
to apex of filament
Dorsilaminar. On dorsal side of blade
Dorsiventral. Planate and having distinct dorsal
and ventral surfaces, the two usually different
Dotted. The color disposed in very small round
spots
Downy. Covered with short, weak, soft trichomes
Drupe. A fleshy fruit with a stony endocarp, as
in Prunus
Drupecetum. An aggregation of drupelets, as in
Rubus
Drupelet. A small drupe, as in Rubus
Dwarf. Very small. Dwarf Shoots or Spurs - Shoots
that develop from preformed buds which have very short internodal
lengths or intervals
Dyad. Pollen grains occurring in clusters of two
Eccentric. Off-center style
Echinate. Covered with spines
Ectocarp
or Exocarp. Outermost layer of pericarp
Edged. One color is surrounded by a very narrow
rim of another
Elaminate. Without blade
Elater. One of four elongate appendages on the
spores, as in Equisetum
Elliptic. With widest axis at midpoint of structure
and with margins symmetrically curved
Emarginate. Lobe rounded; sinus depth 1/16-1/8
distance to midpoint of blade; margins straight or convex
Embryo. Young sporophyte consisting of epicotyl,
hypocotyl, radicle, and one or more cotyledons. Embryo Sac -
Female gametophyte. Linear Embryo - Axial embryo several times
longer than broad, straight, curved or coiled; cotyledons not
expanded; endosperm present or absent
Emergent. With part(s) of plant aerial and part(s)
submersed; rising out of the water above the surface
Emersifolious. With emergent leaves
Endocarp. Innermost differentiated layer of pericarp
Endosperm. Food reserve tissue in seed derived
from fertilized polar nuclei; or food reserve derived from megametophyte
in gymnosperms
Endospermous or Albuminous. With food reserve
in endosperm or albumen, derived from fertilized polar nuclei
Entire. Without indentations or incisions on maigins;
smooth
Entomophily. Pollinated by insects
Eophyll. Term applied to first few leaves with
green, expanded lamina developed by seedlings; transitional
type leaves developed before formation of adult leaves
Epetiolate. Without petiole, leaf sessile
Epetiolulate. Without petiolule, leaflet sessile
Ephemeral. Germinating, growing, flowering and
fruiting in a short period, as most desert herbs
Epicalyx. Group of leaves resembling sepals below
the true calyx
Epicalyx
or Calycle. A whorl of bracts below but resembling
a true calyx
Epicormic Shoots or Water Sprouts. Shoots that
develop from dormant lateral buds on the trunk which have very
long and frequently variable internodal lengths or intervals
Epicotyl. Apical end of embryo axis that gives
rise to shoot system
Epigyny. The condition in which the sepals, petals,
stamens are attached to the floral tube above the ovary with
the ovary adnate to the tube or hypanthium
Epihyperigyny. The condition in which the sepals,
petals, stamens are attached to the floral tube or hypanthium
surrounding the ovary; a combination perigyny and partly inferior
ovary
Epihypogyny. The condition in which the sepals,
petals, stamens are attached about half-way from the base of
the ovary to the partly adnate hypanthium tube; half-inferior
insertion of parts
Epimatium. Fleshy covering of the seed and more
or less fused with the integument; arising from the chalazal
end of the ovule like an additional integument; e.g., Podocarpus
Epiperigyny. The condition in which the sepals,
petals, stamens are attached to the floral or hypanthium cup
above the ovary with the lower part of the hypanthium completely
adnate to the ovary
Epipetalous. With stamens attached to or inserted
upon petals or corolla
Epipetiolar. With branches arising from buds on
the petiole
Epipetric. Upon rock
Epiphyllous. From a phylloclad or peculiar bract,
as in Tilia
Epiphytic. Upon another plant
Epirhizous. With roots upon another plant
Episepalous. With stamens attached or inserted
upon sepals or calyx
Epitropous . Dorsal - Ovule pendulous or hanging, micropyle
above, raphe dorsal (away from ventral bundle). Ventral - Ovule
pendulous or hanging, micropyle above, raphe ventral (toward
ventral bundle)
Equilateral. With halves or sides equal in shape
and size
Equinoctial. Having flowers which expand and close
regularly at particular hours of the day
Equitant. Leaves 2-ranked with overlapping bases,
usually sharply folded along midrib
Eramous. With unbranched stems
Erect. Upright
Erose. Irregularly, shallowly toothed and/or lobed
margins; appearing gnawed
Essential Organs. The androecium and gynoecium
Ethereal
Oil Producing Glands. Mostly aromatic compound
producing glands found on various parts of the plant, without
definitely known functions
Eucamptodromous. With a single primary vein, the
secondary veins curved upward and gradually iminishing distally
within the margin and interconnected by a series of cross-veins
without forming conspicuous marginal loops
Eusporangiate. Having the sporangium develop from
a great amount of leaf tissue as opposed to only one or a few
cells
Eustele. A dissected siphonostele with phloem
only to the outside of the xylem
Evanescent. Passing away, disappearing early
Evergreen. Persistent two or more growing seasons
Evident. Clearly visible macroscopically
Excentric. One-sided; off-center
Excurrent. Running out, as the nerve of a leaf
projecting beyond the margin
Exospore
or Exine. Outer spore wall layer
Exsserted. Projecting out of, beyond
Exstipellate. Without stipels
Exstipulate. Without stipules
Falcate
or Seculate. Sickle-shaped
False
Veins. Small vein-like areas of thick-walled
cells in the leaves of some lower vascular plants
Farinaceous. Mealy
Fasciated. Unnaturally and often monstrously connate
or adnate, the coalesced parts often unnaturally proliferated
in size and/ or number; e.g., inflorescence of Celosia
Fascicle. Cluster of needles borne on a minute
determinate short shoot in the axil of a primary leaf (bract);
e.g. Pinus.
Fascicle
Sheath - Closely imbricated bud scales at the base of the
fascicle of needles; e.g., Pinus
Fascicled. Fleshy or tuberous roots in a cluster
Fasciculate. Leaves or other structures in a cluster
from a common point
Fastigiate. Strictly erect and parallel
Faucal
Area. The throat area
Female,
Carpellate, or Pistillate. Plant with pistillate
flowers only
Fetid. Having an unpleasant, rotten odor
Fenestrate. With windowlike holes through the
leaves or other structures
Fibonacci Phyllotaxis. A fundamental type of leaf
arrangement expressed as a fraction in which each succeeding
fraction is the sum of the two previous numerators and the sum
of the two previous denominators, i.e., 1/2, 1/3, 2/5, 3/8,
5/13, 8/ 21, etc. The numerator represents the number of turns
or spirals around a stem before one leaf is directly above another
and the denominator represents the number of leaves in the turns
or spirals before one is directly above the other. 2/5 phyllotaxy
would mean two twists and five leaves before one leaf is directly
above the other or an angle of divergence of 144 between succeeding
leaves in the stem (2/5 of 360 ).According to Leppik anthotaxis
and semataxis do not necessarily follow the same pattern, with
anthotaxis in Michelia cited as being in 2/7, 3/7, 3/8, and
4/10 systems of arrangement.
Fibrous. With fine, threadlike or slender roots
or fibers
Filament. Stamen stalk
Filiferous. Filamentose or Filiferous With coarse marginal
fibers or threads
Filantherous or Typical. Stamen with distinct
anther and filament with or without thecal appendages, as in
Rhexia or Vaccinium
Filiform. Threadlike, usually flexuous
Fimbriate. Margins fringed
Fimbriolate. Minutely fimbriate
Fistulose. Hollow, as without pith
Flabellate. Fan-shaped branching
Flabelliform. Fan-shaped
Flaccid. Lax and weak
Flagellate. Bearing flagella, whip-like strands
or organs
Fleshy. Succulent roots
Flexuous. With a series of long or open vertical
curves at right angles to the central axis
Floating. Upon the surface of the water
Floccose. Covered with dense, appressed trichomes
in patches or tufts
Flotophyllous. With floating leaves
Flower. Reproductive structure of flowering plants
with or without protective envelopes, the calyx and/or corolla;
short shoot with sporophylls and with or without sterile protective
leaves, the calyx and corolla
Flushing Shoots. Shoots that develop from
mature terminal buds several times during a season. Terminal
bud will develop shoot with new terminal bud which will develop
more shoots and a terminal bud which will develop etc. --several
times in a season with several flushes of growth.
Fly
Trap. Hinged, insectivorous leaf, as in Dionaea
Folded. Foliate embryo with cotyledons usually
thin and extensively expanded and folded in various ways
Follicetum. An aggregation of follicles, as in
Caltha
Follicle. A dry, dehiscent fruit derived from
one carpel that splits along one suture
Fovea. Pit or depression containing the sporangium
in the leaf base of Isoetes
Foveolate. Pitted
Free. Unlike parts or organs unjoined and separate
from one another
Free-adnate. Hypanthium fused with ovary and having
a free limb around or above ovary
Free-central. With the placenta along the central
axis in a compound ovary without septa
Fringe. The modified margin of a petal, sepal,
tepal or lip
Frond. The leaf of a fern
Fruit. Matured ovary of flowering plants, with
or without accessory parts
Fruticose. Shrubby
Fugacious. Ephemeral, usually applied to plant
parts
Funicular. With a persistent elongate funiculus
attached to seed coat, as in Magnolia
Funiculus. Stalk by which ovule is attached to
placenta
Fusiform. Spindle-shaped; broadest in middle and
tapering to each end
Galeate. Helmet-shaped, as one
sepal in Aconitum
Geitonogamy. Fertilization of one flower by another
on the same plant
Gelatinous. Jellylike; soft and quivery
Geminate
or Binate. Paired; in pairs
Gemma. A vegetative reproductive bud borne on
the stem, as in Lycopodium; a multicellular reproductive propagule
on gametophytes, as in ferns
Geniculate. Abruptly bent at a node, zigzag
Geoflorous. With subterranean flower
Geocarpous. Fruits below ground, as in Amphicarpum
Gibbous. Inflated on one side near the base
Gigantic. Very large
Glabrate. Without trichomes
Glabrescent. Becoming glabrous
Glabrous. Smooth; devoid of trichomes
Gland. A secreting part or appendage
Glandular. Covered with minute, blackish to translucent
glands
Glans
(Involucre). Nut subtended by a cupulate, dry
involucre, as in Quercus
Glaucescent. Sparingly or slightly glaucous
Glaucous. Covered with a bloom or smooth, waxy
coating
Globose. Round
Glochidiate. With barbed trichomes, glochids,
usually in tufts
Glomerule. An indeterminate dense cluster of sessile
or subsessile flowers
Glume. Bract, usually occurring in pairs, at the
base of the grass spikelet
Glutinous. Having a shiny, sticky surface
Granular. Finely mealy, covered with small granules
Greasy
or Unctuous. Slick, oily, slippery to touch
Gynagamous. Inflorescence with pistillate flowers
inside or above and neuter flowers outside or below (gynagamocephalous)
Gynandrial or Gynostemial. With fused stamens
and carpels (stigma and style) as in the Orchidaceae
Gynecandrous. Inflorescence with pistillate flowers
inside or above and staminate outside or below, as in spikes
of some species of Carex
Gynehermaphroditic. Inflorescence with pistillate
flowers inside or above and hermaphroditic outside or below
(gynehermaphrodicephalous)
Gynobasic. Attached at base of ovary in central
depression
Gynodioecious. Some plants with perfect flowers
and some with pistillate
Gynoecium or Pistil. The whorl or group of carpels
in the center or at the top of the flower; all carpels in a
flower
Gynomonoecious. Plant with pistillate and perfect
flowers
Gynophore. The stipe of a pistil or carpel
Half-inferior. Other floral organs
attached around ovary with hypanthium adnate to lower half of
ovary
Half-terete. Flat on one side, terete on other;
semicircular in cross section
Haplomorphic. Flowers with parts spirally arranged
at a simple level in a semispheric or hemispheric form; petals
or tepals colored; parts numerous; e.g., Nymphaea, Magnolia
Hastate. Lobe pointed and oriented outward or
divergent in relation to petiole or midrib; sinus depth variable;
margins variable
Hastiform. Triangular with two flaring basal lobes
Haustorial. Absorbing roots, within host of some
parasitic species
Helicoid
Cyme or Bostryx. A determinate inflorescence
in which the branches develop on one side only, appearing simple
Hemianatropous or Hemitropous. With body half-inverted
so that funiculus is attached near middle with micropyle terminal
and at right angles
Herb. A usually low, soft, or coarse plant with
annual aboveground stems
Herbaceous. Soft and succulent
Hermaphrodagamous. Inflorescence with hermaphroditic
flowers inside or above and neuter outside or below (hermaphrodagamocephalous)
Hermaphrodandrous. Inflorescence with hermaphroditic
flowers inside or above and staminate outside or below (hermaphrodandrocephalous)
Hermaphroditic or Monoclinous. Plant with all
flowers perfect
Hesperidium. A thick-skinned septate berry with
the bulk of the fruit derived from glandular hairs, as in Citrus
Heterandrous. With stamens of different sizes
and/or shapes
Heteranthous. Having different states in two different
sets of flowers, only one state present in each set
Heteroblasty. With juvenile foliage distinctly
different from adult foliage in size or shape
Heterocarpous. With carpels of different sizes
and/or shapes
Heterocephalous, Heterocymous, Heterospicous.
Heads, cymes, spikes with flowers of different sexual conditions.
Note: other inflorescence word stems could be used for appropriate
inflorescence type.
Heterocladous. With stems of different sizes and/or
shapes
Heteromerous or Anisomerous. With different number
of members in different whorls
Heteropetalous. With petals of different sizes
and/or shapes
Heterophyllous. With leaves of different sizes
and/or shapes. Heterophyllous Shoots - Shoots that develop from
winter buds which do not contain the primordia of all the leaves
to develop during the year
Heterophytous. Having different states in two
different sets of plants, only one state present in each set
Heterosepalous. With sepals of different sizes
and/or shapes
Heterosexual. Inflorescences or flowers within
the plant with different sexual conditions
Heterosporous. Having two kinds of spores, usually
differing in sizeHeterostichus. With unequal rows
Heterostylous. With styles of different sizes
or lengths or shapes within a species
Heterotropous. Ovule position not fixed in ovary
Hibernal
or Hiemal. Appearing in the winter
Hilum. Funicular scar on seed coat
Hip
or Cynarrhodion (Receptacle and Hypanthium).
An aggregation of achenes surrounded by an urceolate receptacle
and hypanthium, as in Rosa
Hippocrepiform. Horseshoe-shaped
Hirsute. Covered with long, rather stiff trichomes
Hirsutullous or Hirtellous. Minutely hirsute
Hispid. Covered with very long, stiff trichomes
Hispidulous. Approaching hispid, minutely hispid
Homandrous. With stamens of same size and shape
Homanthous. Having more than one state within
each individual flower, all flowers the same
Homocarpous. With carpels of same size and shape
Homocephalous, Homocymous, Homospicous. Heads,
cymes, spikes with flowers sexually uniform
Homochlamydeous. With perianth composed of similar
parts, each part a tepal
Homogamous. With maturation of stamens or anther
and carpels or stigma at same time
Homophytous. Having more than one state within
each individual plant, all plants the same
Homosexual. Inflorescences or flowers sexually
uniform
Homosporous. Having spores of only one kind
Homostylous. With styles of same sizes or lengths
and shapes
Hood. A cover-shaped perianth part, usually with
a turned down margin
Horn. A curved, pointed and hollow protuberance
from the perianth
Hyaline. Thin and translucent or transparent
Hydrophily. Pollinated by water
Hymenopterophily. Pollinated by bees
Hypanepigyny. The condition in which the sepals,
petals, stamens are attached to the elongate floral tube or
hypanthium above the inferior ovary, as in Oenothera
Hypanthium. The fused or coalesced basal portion
of floral parts (sepals, petals, stamens) around the ovary
Hypanthodium. An inflorescence with flowers on
wall of a concave capitulum, as in Ficus
Hyphodromous. With a single primary vein and all
other venation absent, rudimentaryj or concealed within a coriaceous
or fleshy blade
Hypocotyl. Embryonic stem in seed, located below
cotyledons
Hypocotylespermous or Macropodial. With food reserve
stored in hypocotyl, derived from zygote
Hypogynium. Perianth-like structure of bony scales
subtending the ovary, as in Scleria and other members of the
Cyperaceae
Hypogyny. The condition in which the sepals, petals,
stamens are attached below the ovary
Hypophyllous. With small leaves, as bracts, scales,
cataphylls
Hypotropous. Dorsal - Ovule erect, micropyle below,
raphe dorsal (away from ventral bundle). Ventral - Ovule erect,
micropyle below, raphe ventral (toward ventral bundle)
Hysteranthous. With leaves appearing after flowers
Imbricate. Having margins overlapping
Imparipinnate. Pinnate with a conform terminal
leaflet. Imparipinnately Compound - Odd-pinnately compound,
with a terminal leaflet
Imperfect or Unisexual. With stamens or carpels
absent in the flower
Implicate. With both lamina margins folded sharply
inward
Incised. Margins sharply and deeply cut, usually
jaggedly
Inclinate. With lamina folded or curved transversely
near the apex
Inclined. Ascending at 46-75° angle of divergence
Included
Veinlets. Veins ending inside areoles
Incomplete. Leaf without one or more parts: blade,
petiole, stipules; one or more types of floral parts absent
Incrassate. Thickened
Incumbent or Notorhizal. Reclinate with sides
of cotyledons against hypocotyl
Incurved. Curved inward or upward
Indehiscent Capsule. One that does not dehisce
at maturity, as in Peplis
Indeterminate or Evergrowing. Continual growth
of plant parts, not limited by a cessation of meristematic activity
Induplicate. Having margins bent inward and touching
margin of each adjacent structure
Indurate. Hardened
Indusium. A flap of tissue covering a sorus
Inequilateral. With unequal sides
Inermous. Unarmed, without prickles or spines
Inferior. Other floral organs attached above ovary
with hypanthium adnate to ovary
Inflated. Swollen or thickened, as in Eichhornia
Inflexed. Bent abruptly inward or upward
Infrafoliar. On the stem below the leaves, as
in the Arecaceae
Infrapetiolar or Subpetiolar. Axillary bud surrounded
by base of petiole
Infundibular. Funnel-shaped
Integuments. Outer covering of ovule; embryonic
seed coat
Intercalary. Growth region near the base of an
internode or base of blade
Interfoliar. On the stem between the leaves, as
in the Arecaceae
Intermittent. A renewal and cessation of meristematic
activity which produces clusters of stems and/or leaves along
an axis
Internode. A section or region of stem between
nodes
Interpetiolar. With connate stipules from two
opposite leaves
Interrupted or Discontinuous. Symmetry of arrangement
broken, with uneven lengths of internodes
Interstitial. Growth all-over in an organ, no
localized meristems, as in some fruits
Investing. Axial embryo usually erect with thick
cotyledons overlapping and encasing the somewhat dwarfed hypocotyl;
endosperm wanting or limited
Involucel. Small involucre; secondary involucre
Involucre. A group or cluster of bracts subtending
an inflorescence
Involute. Margins or outer portion of sides rolled
inward over upper or ventral surface
Irregular. With floral parts within a whorl dissimilar
in shape and/ or size
Isocotylous. With cotyledons of same size and
shape
Isodynamous. With equally developed structures
Isomerous. With same number of members in different
whorls
Isopetalous. With petals of same size and shape
Isophyllous. With leaves of same size and shape
Isosepalous. With sepals of same size and shape
Isostichous. With equal rows
Jointed. With stems that can be
pulled apart easily at the nodes, as in Equisetum
Keel. The two united petals of
a papilionaceous flower; any structure ridged like the bottom
of a boat
Lacerate. Margins irregularly
cut, appearing torn
Laciniate. Cut into closely parallel ribbonlike
or straplike projections
Lacuna. Chamber or internal air space
Lamina. The leaf tissue other than the veins or
axes
Laminar. Leaf-like stamen without a distinct anther
and filament but with embedded or superficial microsporangia,
as in Degeneria
Laminate. With the placenta over the inner surface
of the ovary wall
Lammas
Shoots. Abnormal late season shoots that develop
from the terminal bud, not a recurring phenomenon as in flushing
shoots
Lanate. Covered with long, intertwined trichomes,
cottony
Lanceolate. Lance shaped, much longer than wide;
widened at or above the base and tapering to the apex
Lanuginose. Cottony, similar to lanate but trichomes
shorter
Lateral
or Axillary. On the side of a structure or at
the nodes of the axis. Lateral Embryo - Basal or baso-lateral
embryo, discoid or lenticular, usually surrounded by copious
endosperm. Lateral Leaf - Leaf on the side of the stem, as in
heterophyllous species of Selaginella
Laterospermous. On the side of the seed
Latiflorous. With broad-flowers
Leaf. A photosynthetic and transpiring organ,
usually developed from leaf primordium in the bud; an expanded,
usually green, organ borne on the stem of a plant. Leaf Primordium
- Meristematic tissue that gives rise to a leaf. Leaf Scar -
A mark indicating former place of attachment of petiole or leaf
base. Linear Leaf - Narrow, flattened, triangular, or quadrangular
leaf usually 1/2-2 in. long; e.g., Taxus, Picea
Leaflet. A distinct and separate segment of a
leaf
Leaf-opposed. On stem opposite the base of the
leaf, as in Alchemilla
Legume. A usually dry, dehiscent fruit derived
from one carpel that splits along two sutures
Lemma. Outer scale subtending grass floret
Lenticel. A pore in the bark
Lenticular. Biconvex, usually elongate and flattish
Lepidote
or Squamulose. Covered with minute scales
Leptophyllous. With leaves to 25 sq. mm in size.
Leptosporangiate. Having the entire sporangium
develop from a periclinal division of a superficial cell or
small group of cells
Ligneous. Woody
Ligulate
or Ray. Strap-shaped
Ligule. An outgrowth or projection from the top
of the sheath, as in the Poaceae; the strap-shaped portion of
a ray or ligulate corolla; a small membranous outgrowth or projection
at the base of the leaf, appearing above the sporangium in fertile
leaves, as in Selaginella and Isoetes
Limb. Expanded portion of corolla or calyx above
the tube, throat or claw
Lineate. In lines, stigmatic surface linear
Lingulate. Tongue-shaped, plano-convex in cross
section
Lip
or Labellum. Either of two variously shaped
parts into which a corolla or calyx is divided, usually into
an upper and lower lip, as in the Lamiaceae and Orchidaceae.
Lip Cells - The line of cells between which the sporangium dehisces
Lobate. Lobe rounded; sinus depth variable; outer
and inner margins concave
Lobe. Any, usually rounded, segment or part of
the perianth
Lobed. Divided into lobes
Locule. Compartment of an anther; ovary cavity
Loculicidal Capsule One that dehisces longitudinally into
the cavity of the locule, as in Epilobium
Lodicule. Abortive perianth part in the Poaceae;
hyaline scales at base of ovary in the Poaceae
Loment. A legume that separates transversely between
seed sections
Long
Shoot. Elongated internodes, rapid annual growth.
Long Bud Shoots - Abnormal buds or shoots which elongate, then
have arrested growth without the development of leaves and lateral
branches
Longitudinal. Dehiscing along long axis of theca
Loose,
Distant, or Scattered. Parts widely separated
from one another, usually irregularly
Lorate. With elongate vertical waves in the margins
or sides at right angles to the longitudinal axis
Lunate. Crescent-shaped, with acute ends
Lyrate. Lyre-shaped; pinnatifid with large terminal
lobe and smaller lower lobes
Major. Greater in size
Malacophily. Pollinated by snails or slugs
Male
or Staminate. Plant with staminate flowers only
Marbled. A surface traversed by irregular veins
of color; as block of marble often is
Marcescent. Usually ephemeral with persistent
remains; withering persistent
Marginal. Pertaining to the border or edge
Massula. A clump of microspores, as in Azolla
Matutinal. With flowers opening in the morning
Medial. Upon or along the longitudinal axis
Median. With stipules adnate to petiole with free
part of stipules near middle of petiole
Median
Leaf. Leaf on top of stem, as in heterophyllous
species of Selaginella
Megasporangium. The sporangium in which megaspores
are produced
Megaspore. A spore that gives rise to a female
gametophyte
Megasporophyll. Modified leaf bearing ovules;
e.g., Zamia
Melittophily. Pollinated by bees
Membranous. Thin and semi-translucent; membrane-like
Meniscoidal. Thin and concaveconvex
Mericarp. A portion of fruit that seemingly matured
as a separate fruit
Mermaphrodigynous. Inflorescence with hermaphroditic
flowers inside or above and pistillate outside or below (hermaphrodigynecephalous)
Mesocarp. Middle layer of pericarp
Metaphyll. Adult leaf
Micro. Axial embryo in minute seeds, less than
0.2 mm long; minute and undifferentiated to almost total size
of seed
Micromelittophily. Pollinated by small bees
Micropyle. Hole through integument(s)
Microsporangium. The sporangium in which microspores
are produced
Microspore. A spore that gives rise to a male
gametophyte
Microsporophyll. Modified leaf bearing microsporangia
or pollen sacs
Midrib. The central conducting and supporting
structure of the blade of a simple leaf
Midvein. The central conducting and supporting
structure of the blade of a leaflet
Minor. Smaller in size
Minute. Very small
Mixed. Contains flower, leaf, and stem primordia;
will give rise to branch with leaves and flower(s)
Monad. Pollen grains occurring singly
Monadelphous. With one group of stamens connate
by their filaments
Moniliform. Elongate roots with regularly arranged
swollen areas
Monocarpellate. One-carpelled
Monocarpic, Hapaxanthic. Perennial or annual,
flowering and fruiting once, then dying; fruiting once
Monocephalous. One-headed, as in composites
Monochasium. A cymose inflorescence with one main
axis
Monocotyledonous. With one cotyledon
Monocyclic. One-whorled
Monoecious. With staminate and carpellate flowers
on same plant
Monoecious. Plant with all flowers imperfect,
but staminate and pistillate flowers on same plant
Monolete
Spore. Bean-shaped, with a single scar line
Monomerous. Whorl with one member
Monomorphic. All of the same shape and size
Monophyllous. One-leaved
Monopodial. Branching with a main axis and reduced
or missing laterals; excurrent
Mucilaginous. Gummy or gelatinous
Mucronate. Less than 3:1 l/w, straight and stiff
Mucronulate. 1:1 l/w or broader than long; straight
Multicellular. Many-celled
Multicipital. With many axes or stems from one
rootstock or caudex
Multicostal. Many-ribbed
Multilocular. Many-locular
Multinodal Shoot. Spring shoot developing from
the terminal winter bud and producing 2 or more whorls of branches;
the cones are partly lateral in the middle of the shoot; e.g.,
Pinus echinata
Multiseriate. Many-rowed; in many series
Multistriate. Many-lined
Muricate. Covered with short, hard protuberances
Muriculate. Minutely muricate
Muticous. Without a vein extension, awn or hair
Myiophily. Pollinated by diptera
Myrmecophily. Pollinated by ants
Nanophyllous. With leaves to 225
sq. mm in size.
Napiform. Turnip-shaped
Navicular. Boat-shaped
Neck. Narrowed portion of hypanthium, between
the base and a flared limb
Necrocoleopterophily. Pollinated by carrion beetles
Nectar
Secreting Glands. Sugary compound producing
glands found mostly on floral parts that produce attractants
for pollinators
Nectary. A specialized nectar-secreting structure
or area
Needle. Acicular; slender, elongated leaf, usually
over 2 in long; e.g., Pinus
Net. Veins uniting to form a network
Neuter
or Agamous. Without stamens and carpels in flower;
or sex organs abortive
Nocturnal. Opening during night
Nodal. At the nodes
Node. Point on the stem where leaves are attached;
or the point of branching of the stem
Nodiform
or Nodulose. Knotty or knobby, as the roots
of most of the Fabaceae
Nucellus. Female sporangium within ovule; megasporangium
in seed plants
Nut. A one-seeded, dry, indehiscent fruit with
a hard pericarp, usually derived from a one-loculed ovary
Nutlet. A small nut
Obconic. Inversely conical
Obcordiform. Inversely cordiform
Obdiplostemonous. With stamens in two whorls,
outer opposite petals, inner opposite the sepals
Oblique. Having an asymmetrical base
Oblong. With widest axis at midpoint of structure
and with margins essentially parallel
Obovate. Inversely ovate
Obscure. Not clearly visible macroscopically,
usually owing to incomplete differentiation
Obtriangular. Inversely triangular
Obtrullate. Inversely trullate
Obtuse. Margins straight to convex, forming a
terminal angle more than 90
Ocellated. A broad spot of some color has another
spot of a different color within it
Ocreate. Having a stipular tube surrounding stem
above insertion of petiole or blade
Ocreolate. Diminutive of ocreate; usually applied
to bract bases
Oligomerous. With reduction in number of members
within whorl
Oligotaxy. Reduction in number of whorls
Opaque. With a dull surface
Operculate Capsule. One that dehisces through
pores, each of which is covered by a flap, cap, or lid, as in
Papaver
Opposite. Two leaves or other structures per node,
on opposite sides of stem or central axis
Ornithophily. Pollinated by birds
Orthotropous or Atropous. With straight body so
that funicular attachment is at one end and micropyle at other
Osseous. Bony
Ovary. Ovule-bearing part of pistil
Ovate. With widest axis below middle and with
margins symmetrically curved; egg-shaped
Ovulate. Female; bearing ovules only in the flowers
Ovule. Embryonic seed consisting of integument(s)
and nucellus
Ovuliferous Scale. Highly modified lateral branch
in the axil of a leaf (bract), and bearing ovules; may be flat
or peltate, woody or fleshy; e.g., Pinaceae
Pachycauly. Short, thick, frequently
succulent stems, as in
Painted. Colors disposed in streaks of unequal
intensity
Palate. The raised area in the throat of a sympetalous
corolla
Palea. Inner scale subtending grass floret
Paleaceous. With small membranous scales, chaffy
Palinactinodromous. Actinodromous, the primary
veins with one or more subsidiary radiations above the primary
one
Palmate. Radiately lobed or divided. Palmately
Compound - With leaflets from one point at end of petiole
Palmate-pinnate. With first order leaflets palmately
arranged, second order pinnately arranged
Palmatifid. Cut palmately
Palmatisect. Sectioned or divided palmately into
distinct segments
Panduriform. Fiddle-shaped; obovate with sinus
or indentation on each side near base and with two small basal
lobes
Panicle. Branched inflorescence with pedicelled
flowers
Pannose
or Felted. With matted, feltlike layer of trichomes
Papilionaceous. With large posterior petal (banner
or standard) two lateral petals (wings) and usually two connate
lower petals (keel); as in the Fabaceae
Papillose. Covered with minute tubercles
Pappus. Bristly or scaly calyx in the Asteraceae
Parallel. Thecae or anther cells along side of
the connective or longitudinal to each other; with veins extending
from base to apex, essentially parallel
Parallelodromous. With two or more primary veins
originating beside one another at the blade base and running
more or less parallel to the apex where they converge
Paraphyses. Hairs or hair-like structures in the
sorus
Parietal. With the placentae on the wall or intruding
partitions of a unilocular compound ovary
Paripinnately Compound. Even-pinnately compound,
without a terminal leaflet
Parted. Indentations or incisions cut 1/2-3/4
distance to midrib
Patelliform. Knee-shaped; disk-shaped
Patent. Spreading
Pectinate. Pinnatifid with closely set segments;
comb-like
Pedate. Palmately cut or divided with the lower
pair basiscopically exaggerated
Pedicel. Individual flower stalk
Peduncle. Main stalk for entire inflorescence
Peg
(sterigmata). Lateral stem projection to which
leaf is attached and persistent after leaf dehiscence; ie.,
abscission layer between peg and leaf; leaf may be sessile;
e.g., Picea; or petiolate; e.g. Tsuga, on the peg
Pellucid. Clear, transparent
Peltate. Usually having petiole attached near
the center on the underside of blade
Peltiform. Rounded with petiole attached to center
of blade or apparently to laminar tissue
Pendent. Drooping, hanging down; pendulous
Pendulous. Hanging loosely or freely; pendent
Penni-parallel. With veins extending from midrib
to margins, essentially parallel
Pentacyclic. Five-whorled
Pentagonal. Five-angled
Pentamerous. Whorl with five members
Pentandrous. With five stamens
Pepo. A berry with a leathery nonseptate rind
derived from an inferior ovary, as in Cucurbita
Perennial, Polycarpic. Living more than two years;
fruiting more than once
Perfect
or Bisexual. With both stamens and carpels or
pistils in the flower
Perfoliate. Having base completely surrounding
stem
Perianth. An aggregation of tepals or combined
calyx and corolla
Pericarp. Fruit wall
Pericarpous. Around the fruit
Pericladial. With a sheathing base, as in the
Apiaceae
Perigynium. Sac-like bract subtending the pistillate
flower, as in Carex
Perigyny. The condition in which the sepals, petals,
stamens are attached to the floral tube or hypanthium surrounding
the ovary with the tube or hypanthium free from the ovary
Peripheral. On the outer surface or edge
Perisperm. Food reserve in seed derived from diploid
nucellus or integuments
Perispermous. With food reserve in perisperm,
derived from diploid nucellus or integuments
Perispore or Perine. An outer covering of some
fern spores, with different configuration than that of the exospore
Persistent. Remaining attached; applied to individual
parts
Personate. Two-lipped with the upper arched and
the lower protruding into corolla throat
Petal. A corolla member or segment; a unit of
the corolla
Petalantherous. With a terminal anther and distinctly
petaloid filament, as in Saxifraga
Petaliferous. Most frequently at the base of petals
Petaloid. Petal-like in shape, texture and/or
color
Petalostemonous. With filaments fused to corolla,
anthers free
Petiolar. May be all over (general) or near apex
(acropetiolar) or near base (basipetiolar)
Petiolate. With a petiole
Petiole. Leaf stalk
Petiolulate. With a petiolule
Petiolule. Leaflet stalk
Petrorhizous. With roots on rock
Phalaenophily. Pollinated by moths
Phanerantherous. With stamens exserted
Phanerocotylar or Epigeous. With the cotyledons
emergent from seed, usually appearing above ground
Photosynthetic. Blade-like and green
Phyllary. One of the involucral leaves subtending
a capitulum, as in composites
Phyllodial. Flattened and blade-like
Phyllodium. Flattened blade-like petiole or midrib
Phyllopodic. With blade-bearing leaves at base
of plant
Phyllotaxis. Arrangement of leaves, primarily
photosynthetic in function
Piliferous. More than 20:1 l/w, hair-like, flexuous
Pilose. With soft, shaggy trichomes
Pinna. A primary division of a fern leaf
Pinnate. Compound, with the leaflets arranged
on both sides of a common axis. Interruptedly Pinnately Compound
- With smaller and larger leaflets alternating along the rachis
Pinnate-pinnatifid. Pinnate with pinnatifid pinnae
Pinnatifid. Pinnately cut, more than half way
to the midvein
Pinnatisect. Sectioned or divided pinnately into
distinct segments
Pinnule. A secondary division of a fern leaf
Pisiform. Pea-shaped
Pistil. One or more fused carpels consisting of
stigma, style (if present) and ovary
Pistillate, Carpellate, or Female. With pistils
or carpels only in the flower
Pitcher. Ventricose to tubular insectivorous leaf,
as in Sarracenia
Pith. Centermost tissue of stem, usually soft
Placenta. Ovule-bearing region of ovary wall
Planate
or Plain. With lamina flat, without folds or
rolls
Platycanthous. With flat and usually large spines
Plectostele. A protostele dissected into anastomosing
plate-like units
Pleiochasium. Compound dichasium in which each
cymule has three lateral branches
Pleiomerous. With increase in number of members
within whorl
Pleiotaxy. Increase in number of whorls
Pleomorphic. Actinomorphic with numbers of parts
reduced; e.g., Tripogandra
Pleurogamy. Pollen tube entrance through side
of ovule
Pleurotropous. Dorsal - Ovule horizontal, micropyle
toward ventral bundle, raphe above. Ventral - Ovule horizontal,
micropyle toward ventral bundle, raphe below
Plicate
or Plaited. Fluted, longitudinally folded
Pliestesial or Multiperennial. Monocarpic but
living several to many years before flowering, as in Agave
Plumose. Feather-like
Plumule. Embryonic leaves in seed derived from
epicotyl
Pneumatophorous. With spongy, aerating roots,
usually found in marsh plants
Pollen
Grain. Young male gametophyte. Pollen Sac -
Male sporangium
Pollinia. Grains occurring in uniform coherent
masses
Polyad. Pollen grains in clusters of more than
four
Polyandrous. Many-stamened
Polyanthous. Having different states in several
to many (more than three) different sets of flowers, only one
state present in each set
Polycarpellate. Many-carpellate
Polycephalous. Many-headed, as in composites
Polycyclic. Many-whorled
Polydelphous. With several groups of stamens connate
by their filaments
Polygamo-dioecious. Plants dioecious, but with
some perfect flowers on staminate or pistillate plants or both
Polygamo-monoecious. Plant monoecious, but with
some perfect flowers
Polygamous. Plant with perfect and imperfect flowers
Polyheterophytous. Having different states in
several to many (more than three) different individuals or sets
of plants, only one state present in each set
Polymerous. Whorl with many members
Polymorphic. Having several to many (more than
three) different shapes and/or sizes within the same species
Polyphore. A receptacle or torus bearing many
distinct carpels, as in Rosa
Polystichous. Leaves or other structures in many
rows
Pome
(Receptacle and Hypanthium). A berry-like fruit,
adnate to a fleshy receptacle, with cartilaginous endocarp,
as in Malus
Poricidal. Dehiscing through a pore at apex of
theca
Porogamy. Pollen tube entrance through the micropyle
Posterior. Lobe - The lobe next to axis, away
from the subtending bract; adaxial lobe. Posterior Ridges, Lines,
Grooves - The lines, grooves, ridges in or on the ventral side,
adaxial, within the perianth
Pouch
or Sac. A bag-shaped structure
Precocious. Developing unusually early
Preformed Shoots. Normal shoots that develop from
winter buds which contain primordia of all leaves that will
expand during the season
Prickle. A sharp pointed outgrowth from the epidermis
or cortex of any organ
Prickly. With prickles
Primary. From radicle of embryo; tip of main axis
Primocane. The first year non-flowering stem,
as in most blackberries; a turion
Primordium. Cells in embryo or bud giving rise
to roots, leaves or flowers. Protected Primordium - Shoot and/or
flower primordia surrounded by scales. Naked Primordium - Shoot
and/or flower primordia not surrounded by scales
b. Flowering before normal period, as spring flowers
in the fall
Procumbent, Prostrate, or Reclining. Trailing
or lying flat, not rooting at the nodes; humistrate
Proleptic Shoots. Abnormal late season shoots
that develop from the lateral buds immediately beneath the terminal
Promeristem. Apical growing or meristematic tissue
that gives rise to other bud parts
Prop
or Stilt. Adventitious, supporting roots usually
arising at lower nodes
Prostrate. Flat, spreading; growing low along
the ground
Protandrous. With stamens or anthers developing
before carpels or stigma
Protantherous. With leaves appearing before flowers
Prothallus. Gametophyte of lower vascular plants
Protogynous. With carpels or stigma maturing before
stamens or anthersProtostele. Stele having a solid
column of vascular tissue with xylem centrally located
Proximal. Near the point of origin or attachment
Pruinose, Frosted, or Sebiferous. With a heavy
wax coat
Pseudanthium. Several flowers simulating a simple
flower but composed of more than a single axis with subsidiary
flowers
Pseudocarp (Receptacle). An aggregation of achenes
embedded in a fleshy receptacle, as in Fragaria
Pseudodrupe (Involucre). Two-four loculed nut
surrounded by a fleshy involucre, as in Juglans
Pseudomonomerous. Whorl seemingly with one member
which is a fusion product of two or more parts
Pseudo-terminal. Bud appearing apical but is lateral
near apex, developing with death or nondevelopment of terminal
bud
Psychophily. Pollinated by butterflies
Pterocauly. Winged stems
Puberulent. Minutely pubescent
Pubescent. Covered with dense or scattered trichomes
Pulverulent. Covered with fine, powdery wax granules
Pulvinal. With a swollen base, as in the Fabaceae
Pulvinus. The swollen base of a petiole or petiolule
Punctate. Covered with minute impressions or depressions
Pungent. Acrid; terminating in a rigid sharp point
Purcate. Forked
Pustulate. With scattered blisterlike swellings
Pyrene. Fleshy fruit with each seed surrounded
by a bony endocarp, as in Ilex
Pyriform. Pear-shaped
Quadrate. Nearly square to form
Quadrifid. Cut or divided into four lobes or parts
Quincuncial. Having five structures, two of which
are exterior, two interior, and a fifth with one margin covering
interior structure and other margin covered by that of one of
the exterior structures
Raceme. Unbranched, indeterminate
inflorescence with pedicelled flowers
Rachilla. Secondary axis of compound leaf; central
axis of a grass or sedge spikelet
Rachis. The main axis of a pinnately compound
leaf; major axis within an inflorescence; axis of a compound
fern blade
Radical. Leaves basal, near ground, usually from
caudex or rootstock
Radicle. Basal end of embryo axis that gives rise
to root system
Rameal
Sheath. Leaf sheath on the stem joints, as in
Equisetum
Ramentaceous. Having many thin scales, as on the
epidermis of some ferns
Ramose. Branched
Raphe. Ridge on seed coat formed from adnate funiculus
Ray. Secondary axis in a compound inflorescence
Receptacle or Torus. The region at end of pedicel
or on axis to which flower is attached; point on a leaf where
sporangia are attached
Receptaculum. A fleshy structure below the seed
formed from the bases of bracts and the swollen receptacle or
cone axis; e. g., Acmopyle, and some Podocarpus spp
Reclinate. Bent down upon the axis, no angle of
divergence
Reclined. Descending at 106-135° angle of divergence
Rectangular. Box-shaped, longer than wide
Recurved. Curved outward or downward
Reduced. Decreased in size
Reflexed. Bent or turned downward
Regular. With floral parts within a whorl similar
in shape and size
Reniform. Kidney-shaped, with shallow sinus and
widely rounded margins
Repand. Sinuate with indentions less than 1/16
distance to midrib or midvein
Repent. Creeping or lying flat and rooting at
the nodes
Replicate. With lamina folded once abaxially along
midrib or midvein
Replum. Persistent septum after dehiscence of
fruits, as in the Brassicaceae
Resinous. Having a yellowish, sticky, exudate
Resupinate. Inverted or twisted 180°, as in pedicels
in the Orchidaceae
Reticulate. Netted; with veins forming a network
Reticulodromous. With a single primary vein, the
secondary veins not terminating at the margin and losing their
identities near the margin by repeated branching, yielding a
dense reticulum
Retinaculum, Jaculator or Echma. A persistent
indurated, hook-like funiculus in the fruits of Acanthaceae
Retrorse. Bent or directed downward. Retrorsely
Crenate - Rounded teeth directed toward base. Retrorsely Serrate
- Sharp or pointed teeth directed toward base
Retuse. Lobe rounded; sinus depth to 1/16 distance
to midpoint of blade; margins convex
Revolute. Margins or outer portion of sides rolled
outward or downward over lower or dorsal surface
Rhizocarpic. Plants having the roots living for
many years with the stems dying annually
Rhizoid. A hair-like absorptive organ on gametophytes
and rarely on sporophytes
Rhizome. A horizontal underground stem
Rhizotaxis. Arrangement of roots
Rhombic. With widest axis at midpoint of structure,
and with straight margins; elliptic but margins straight and
middle angled
Ribbed. With longitudinal nerves
Ringed. With old bud scale scar rings
Robust. Large
Rolled. Sides enrolled, usually loosely, over
upper or lower surfaces
Root. An absorbing and anchoring organ, usually
initially developed from the radicle and growing downward. Root
Cap - Parenchymatous, protective apex of root. Root Hair - Lateral,
absorbing outgrowth of the epidermal cell
Secondary Root. Lateral root with
root cap and hairs, derived from the pericycle
Rootstock. A term applied to miscellaneous types
of underground stems or parts
Roridulate or Dewy. Covered with waxy platelets,
appearing dewy
Rostellum or Beak. Persistent stylar base on fruit
Rosulate. Leaves in a rosette
Rotate. Wheel-shaped, with short tube and wide
limb at right angles to tube
Rounded. Margins and apex forming a smooth arc
Rudimentary. Basal, small nonperipheral embryo
in small to large seed; relatively undifferentiated; endosperm
copious
Rugose. Covered with coarse reticulate lines
Ruminate. Coarsely wrinkled, appearing as chewed
Runcinate. Oblanceolate with lacerate to parted
margins
Runner
or Stolon. An indeterminate, elongate, above
ground propagative stem, with long internodes, rooting at the
tip forming new plants
Saccate. Pouch-like
Sagittate. Lobe pointed and oriented downward
or inward in relation to petiole or midrib; sinus depth variable;
margins variable
Sagittiform. Triangular-ovate with two straight
or slightly incurved basal lobes
Salient,
Porrect, or Projected. Pointed outward, usually
said of teeth
Salverform. Trumpet-shaped; with slender tube
and limb nearly at right angles to tube
Samara. A winged, dry fruit
Samaracetum. An aggregation of samaras, as in
Liriodendron
Sapromyiophily. Pollinated by carrion or dung
flies
Sarcocauly. Fleshy stems
Sarcous. With the seed coat fleshy
Scaberulent. Approaching scabrous
Scabridulous. Minutely scabrous
Scabrous. Having a harsh surface
Scale. Small, non-green leaf on bud and modified
stem; small, scarious to coriaceous flattened bodies within
the perianth, as in the Cyperaceae and Asteraceae. Scale Leaf
- Small, usually appressed and imbricate; e.g., Juniperus, Thuja
Scape. A naked flowering stem with or without
a few scale leaves, arising from an underground stem
Scapose. With a solitary flower on a leafless
peduncle or scape, usually arising from a basal rosette
Scarious. Thin and dry, appearing shriveled
Scarred. With old leaf base, stipular and/or branch
scar regions
Schizocarp. Separated body, as in separating fruits
(achenes, berries, carcerules, follicules, mericarps, nutlets,
samaras), splitting apart at maturity
Sclerocauly. Hard, dryish stems
Sclerous. Hard
Scurfy
or Lentiginous With exfoliating scaly incrustations
Seasonally. Occurring during a seasonal cycle,
or each season
Secund
or Unilateral. Flowers or other structures on
one side of axis
Seed. A matured ovule. Seed Coat - Outer protective
covering of seed
Segment. The ultimate division or unit of a dissected
fern leaf
Semataxis. Arrangement of semaphylls (petals,
sepals, tepals), primarily advertising (pollinator attracting)
in function
Semicarpous. With ovaries of adjacent carpels
partly fused, stigmas and styles separate
Semicraspedodromous. With a single primary vein,
the secondary veins branching just within the margin, one branch
from each terminating at the margin and the other forming a
marginal loop and joining the superadjacent secondary vein
Sepal. A calyx member or segment; a unit of the
calyx
Sepaliferous. Most frequently at the base of sepals
Sepaloid. Sepal-like in shape, texture and/or
color
Septal. At the junction of the septa in the ovary
Septate. Divided by internal partitions into locules
or cells
Septum
or Dissepiment. Partition. Septicidal Capsule
- One that dehisces longitudinally through the septa, as in
Penstemon
Sericeous. With long, silky trichomes, usually
appressed
Serotinal. Opening late; appearing in late summer
Serrate. Saw-toothed; teeth sharp and ascending,
but cut 1/16-1/8 distance to midrib or midvein
Serrated. Cut into sawlike teeth
Serrulate. Diminutive of serrate, but cut to 1/16
distance to midrib or midvein
Sessile. Without a petiole or petiolule
Seta. A hair-like extension of the leaf, as in
homophyllous species of Selaginella
Setose,
Setaceous. Having setae or bristlelike trichomes
Sheath. Any more or less tubular portion of the
leaf surrounding the stem or culm, as in the Poaceae
Sheathing. Having tubular structure enclosing
stem below apparent insertion of blade or petiole
Shining,
Nitid, or Laevigate. Lustrous, polished
Shrub. A much-branched woody perennial plant usually
without a single trunk
Silicle. A dry, dehiscent fruit derived from two
or more carpels that dehisce along two sutures and which has
a persistent partition after dehiscence and is as broad as,
or broader, than long
Silique. A silicle type fruit that is longer than
broad
Simple. Not composed of more than one anatomically
or morphologically equivalent unit
Sinistrorse. Rising helically from left to right,
a characteristic of twining stems
Sinuate. Long horizontal curves in the body of
the structure parallel to the central axis
Siphonostele. A stele having vascular tissue in
the form of a hollow cylinder, with a central pith
Soboliferous. With loosely clumped shoots arising
some distance apart from rhizomes or under ground suckers
Solenostele. A siphonostele having phloem both
internal and external to the xylem
Solitary. One-flowered, not an inflorescence
Sorosis. Fruits on a common axis that are usually
coalesced and derived from the ovaries of several flowers, as
in Morus
Sorus. A cluster of sporangia
Spadix. Unbranched, indeterminate inflorescence
with flowers embedded in the rachis
Spathe. An enlarged bract enclosing an inflorescence
Spathulate or Spatulate. Oblong or obovate apically
with a long attenuate base
Spatulate. Foliate, erect embryo with variable
cotyledons, thin to thick and slightly expanded to broad
Spherical. With multi-dimensional radial symmetry
Sphingophily. Pollinated by hawk moths and nocturnal
lepidoptera
Spiculate. With crystals in or on the surface
Spike. Unbranched, indeterminate, elongate inflorescence
with sessile flowers
Spikelet
or Locusta. A small spike; the basic inflorescence
unit in grasses and sedges
Spine. Sharp-pointed petiole, midrib, vein, or
stipule
Spinose
or Pungent. Acuminate but coriaceous and stiff
Spiral. Twisted like a corkscrew
Spirolobal. With incumbent cotyledons folded once
Spongy. Cellular; sponge-like
Sporangiophore. The umbrella-shaped sporangium-bearing
unit of the strobilus, as in Equisetum
Sporangium. A spore case
Sporocarp. A hard, nut-like structure containing
the sporangia in heterosporous ferns
Sporophyll. A spore bearing leaf
Spotted. The color disposed in small spots
Spur. A short shoot on which flowers and fruits
or leaves are borne; a tubular or pointed projection from the
perianth
Squamose. Having coarse scales
Squarrose. Usually sharply curved downward or
outward in the apical region, as the bracts of some species
of Aster
Stamen. Male sporophyll within the flower; floral
organ that bears pollen in angiosperms
Staminal. Most frequently at base of filament
Staminate or Male. With stamens only in the flower
Staminodial. Frequently entire structure is nectariferous
Staminodium. Sterile stamen, may be modified as
a nectary or petaloid structure. Staminal Disc - A fleshy, elevated
cushion formed from coalesced staminodia or nectaries
Standard, Banner, or Vexillum. The upper, usually
wide petal in a papilionaceous corolla
Stele. The central primary vascular system of
the stem and associated tissues delimited from the cortex by
endodermis and pericycle, Delimited from the cortex by endodermis
and pericycle
Stellate. Star-shaped
Stem. A supporting and conducting organ usually
developed initially from the epicotyl and growing upward
Stereomorphic. Flowers 3-dimensional with basically
radial symmetry; parts many or reduced, and usually regular;
e.g., Narcissus, Aquilegia
Stigma. Pollen-receptive portion of pistil
Stipe,
Podogyne, Carpopodium. Basal stalk. Stipe Bundles
- The vascular bundles of the fern petiole
Stipellate. With stipels
Stipels. Paired scales, spines, or glands at the
base of petiolule
Stipulate. With stipules
Stipules. Paired scales, spines, glands, or blade-like
structures at the base of a petiole. Stipular Scar - A mark
indicating former place of attachment of stipule
Stolon. Runner or indeterminate, elongate, above
ground propagative stem, rooting at the tip producing new plants.
Underground Stolon - A determinate, elongate, underground propagative
stem with long internodes forming a bulb or tuber at the tip
Stoloniferous. Bearing stolons; sarmentose
Stomate. Opening or pore in leaf epidermis; intercellular
space between two guard cells
Stomium. Lip cell region of a fern sporangium
Striate. With longitudinal lines
Strict. Stiff and rigid
Strigillose. Diminutive of strigose
Strigose. Covered with sharp, coarse, bent trichomes
usually with a bulbous base
Striped. Longitudinal stripes of one color crossing
another
Strobilus. Stem with short internodes and spore-bearing
appendages; a cone
Strombiform. Elongate snail-shaped
Strophiolate. With elongate aril or strophiole
in the hilum region
Style. Attenuated, non-ovule-bearing portion of
pistil between stigma and ovary
Stylocarpellous. With a style and without a stipe,
the normal carpel
Stylocarpepodic. With a style and stipe
Stylopodic. With a stylopodium or discoid base,
as in the Apiaceae
Subacrocaulous. With branches at or near tip of
main stem
Subapical. At one side near apex of ovary
Subbasal. Near the base
Subbasicaulous. With branches at or near base
of main stem
Subbasifixed. Anther attached near its base to
apex of filament
Suberous. Corky
Subglabrate. Almost glabrous
Subglobose. Almost round or spherical
Submersed. Beneath the surface of the water
Submersicaulous. With submersed stems
Subtend. To stand below and close to
Subterminal. Near the apex
Subterranean or Hypogeous. Below the surface of
the ground
Subulate. Very narrow and tapering; awl-shaped;
linear
Sucker
Shoots. Shoots that develop from adventitious
buds on old stumps or roots, usually after cutting or injury,
which have elongate internodal lengths and intervals
Suffrutescent. Woody basally, herbaceous apically
Sulcate. With longitudinal grooves
Summer
Annual. Germinating in spring or early summer
and flowering and fruiting in late summer or early fall, then
dying
Superior. Other floral organs attached below ovary
Supervolute. With lamina with one edge tightly
enrolled and with the other loosely enrolled covering the first,
loosely convolute
Supine. Prostrate, with parts oriented upward
Suprafoliar. On the stem above the leaves, as
in the Arecaceae
Suprarhizous. On top of the root
Surcarpous. With fruits on surface of ground
Surcurrent. Extending along stem upward from leaf
base
Surficial or Epigeous. Upon or spread over the
surface of the ground
Syconium. A syncarp with the achenes borne on
the inside of a hollowed-out receptacle or peduncle, as in Ficus
Syleptic
Shoots. Abnormal shoots that develop from lateral
buds before they have reached maturity
Sympetalous. With fused petals
Sympatric. Collocated, together
Sympodial. Branching without a main axis but with
many, more or less, equal laterals
Synantherous. With leaves and flowers appearing
at same time
Syncarpous. With fused carpels
Syncotyly. Cotyledons coalesced, forming a funnel
or trumpet
Syngenesious. With fused anthers
Synovarious. With ovaries of adjacent carpels
completely fused, styles and stigmas separate
Synsepalous. With fused sepals
Synstylovarious. With ovaries and styles of adjacent
carpels completely fused, stigmas separate
Syntropous. With radicle pointing toward hilum
Tannin
Bearing Glands. Tannin producing glands found
in various parts of the plant, presumably protective in some
structures
Tap. Persistent, well-developed primary root
Tendril. Long, slender, coiling branch, adapted
for climbing (most tendrils are leaf structures)
Tentacular. Glandular-haired or tentacle-bearing
insectivorous leaf, as in Drosera
Tepal. A member or segment of perianth in which
the parts are not differentiated into distinct sepals and petals
Terete. Cylindrical and elongate
Tergeminate. With three orders of leaflets, each
bifoliolate, or with geminate leaflets ternately compound
Terminal. At apex or end of stem. Terminal Bud
Scale Scar Rings - Several marks in a ring indicating former
places of attachment of bud scales
Ternately Compound. With leaflets in three's
Tessellated. Color arranged in small squares,
so as to have some resemblance to a checkered pavement
Testa. The outer, commonly hard and brittle seed-coat
Tetracyclic. Four-whorled
Tetrad. Pollen grains in clusters of four
Tetradynamous. With stamens in two groups, usually
four long and two short
Tetragonal. Four-angled
Tetrahedral. Having the form of a tetrahedron
Tetralocular. Four-locular
Tetramerous. Whorl with four members
Tetrandrous. With four stamens
Tetrastichous. Leaves or other structures in four
rows
Theca. One half of anther containing two pollen
sacs or male sporangia
Thorn. A sharp-pointed branch
Throat. An open, expanded tube in the perianth
Thyrse. A many-flowered inflorencence with an
indeterminate central axis and with many opposite lateral dichasia
Tiller. A grass shoot produced from the base of
the stem
Tomentose. Covered with dense, interwoven trichomes
Torose. Cylindrical with contractions at intervals
Tortuous. Irregularly twisted
Trailing. Sprawling on ground, usually with adventitious
roots
Transverse or Explanate. At rightangles to long axis
Tree. A tall, woody perennial plant usually with
a single trunk
Triangular. With three sides and three angles
Tricarpellate. Three-carpellate
Trichome. A leaf or stem surface hair
Tricyclic. Three-whorled
Tridynamous. With stamens in two equal groups
of three
Trifid. Cut or divided into three lobes or parts
Triflorous. Three-flowered
Trifoliate. Three-leaved
Trifoliolate. With three leaflets
Trifurcate. Divided into three forks or branches;
three-forked
Trigonous. Three-angled
Triheteranthous. Having different states in three
different sets of flowers, only one state present in each set
Triheterophytous. Having different states in three
different sets of plants, only one state present in each set
Trilete
Spore. Basically tetrahedral, but often appearing
round or triangular, with three scar lines forming a Y
Trimerous. Whorl with three members
Trimorphic. Having three different shapes and/or
sizes within the same species
Trioecious. Plants staminate, pistillate or perfect
Tripalmately Compound. With three orders of leaflets,
each palmately compound
Tripinnately Compound. With three orders of leaflets,
each pinnately compound
Triquetrous. Three-angled with the sides usually
concave
Tristichous. Leaves or other structures in three
rows
Triternate. With three orders of leaflets, each
ternately compound
Trullate. With widest axis below middle and with
straight margins; ovate but margins straight and angled below
middle, trowel-shaped
Truncate. Cut straight across; ending abruptly
almost at right angles to midrib or midvein
Tryma
(Involucre). Two-four loculed nut surrounded
by a dehiscent involucre at maturity, as in most species of
Carya
Tube. Cylindrical part
Tuber. A thick storage stem, usually not upright
Tuberculate. With hard, swollen, persistent base
or tubercle. Tuberculate or Verrucose - With a warty surface
Tubercules. Silica deposits on the stem ridges,
as in Equisetum
Tuberous. Fleshy roots resembling stem tubers
Tubular. Cylindrical
Turbinate. Top-shaped; obconic
Turgid. Tumid or swollen
Turion. An over-wintering bud, as in Lemna
Twining. Twisted around a central axis
Umbel. A determinate or indeterminate
flat-topped or convex inflorescence with the pedicels arising
at a common point. Compound Umbel - An umbel with primary rays
or peduncles arising at a common point with a secondary umbel
arising from the tip of the primary rays; a branched umbel
Umbellet. The secondary umbel in a compound umbel
Umbilicate. Depressed in the center
Umbo. Projection, with or without spine or prickle,
on the apophysis of the cone scale
Umbonate. Round with a projection in center
Umbraculate. Umbrella-shaped, as in Sarracenia
Umbraculiform. Umbrella-shaped
Undulate. With a series of vertical curves at
right angles to the central axis
Unguiculate. Clawed
Unicarpellous or Stylodious. With solitary, free
carpel in gynoecium
Unifoliolate. With a single leaflet with a petiolule
distinct from the petiole of the whole leaf, as in Cercis
Unilocular. One-locular
Uninodal
Shoot. Spring shoot developing from the terminal
winter bud and producing only one internode with one whorl of
branches at the end; the cones are subterminal at the end of
the shoot; e.g., Pinus resinosa
Uniseriate. One-rowed; in one series
Unisexual. With only one sex in each flower
Urceolate. Urn-shaped
Stinging
Urent. With erect, usually long trichomes that
produce irritation when touched
Utricle. A small, bladdery or inflated, one-seeded,
dry fruit.
Vallecular Canal. A canal beneath
a stem groove
Valvate. Sides enrolled, adaxially or abaxially
so that margins touch
Valvular. Dehiscing through a pore covered by
a flap of tissue. Valvular or Septifragal Capsule - One with
valves breaking away from the septa, as in Ipomoea
Variegated. The color disposed in various irregular,
sinuous, spaces
Vascular
Bundle or Trace Scar A mark indicating former place of attachment
within the leaf scar of the vascular bundle or trace
Vegetative Frond. Frond lacking sporangia
Velum. The membranous flap covering the sporangium,
as in Isoetes
Velutinous. Covered with dense, straight, long
and soft trichomes; pile-like
Ventral. Pertaining to the surface nearest the
axis; inner face of an organ; the upper surface of the leaf;
adaxial. Ventral Side - Top side or upper side of a perianth
part
Ventricose. Inflated on one side near the middle
Ventristipular. On ventral side of stipule
Vermiform. Worm-shaped
Vernal. Appearing in spring
Verrucose. Warty
Versatile. Dorsifixed but anther seemingly swinging
free on the filament
Verticillaster. Whorled dichasia at the nodes
of an elongate rachis
Vespertine. With flowers opening in the evening
or night; appearing or expanding in the evening
Vestigial. Minute; a remnant
Vexillate. Having one structure larger than others
which is folded over smaller enclosed structures
Villosulous. Minutely villous
Villous. Covered with long, soft, crooked trichomes
Vine
or Liana. An elongate, weak-stemmed, often climbing
annual or perennial plant, with herbaceous or woody texture
Virgate. Wand-like; long, slender, and straight
Viscid. Sticky or glutinous
Whorl. A cyclic or acyclic group
of sepals, or petals, or stamens, or carpels
Whorled,
Radiate, or Verticillate. Three or more leaves
or other structures per node
Wing. Lateral petals, as in the Fabaceae; a flattened
extension, appendage or projection from a perianth part
Winged. With flattened blade-like margins. Winged Nut
(Bract) - Nut enclosed in a winglike bract, as in Carpinus
Wood. Xylem consisting of vessels and/or tracheids, fibers,
and parenchyma cells
Woody. Hard and lignified
Zoned. The same as ocellated, but the
concentric bands more numerous
Zonocaulous. With branches intermittently spaced along
main stem
Zygomorphic or Bilateral. With floral parts in two symmetrical
halves
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